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STONY CORALS - Large Polyp Stony and Small-Polyp Stony  and SOFT CORALS

   
    The stony corals all produce a hard skeleton composed of calcium carbonate, a fact reflected in the common name. Accurate identification of stony coral species may pose difficulties even for experts, but most of those available in the saltwater aquarium trade are readily recognizable by genus name. All stony corals require excellent water quality, and all, except one, require bright, broad spectrum lighting. Tubastrea, the Orange Polyp Coral, lacks zooxanthellae and must be maintained in a dimly lit  saltwater aquarium. It also requires feeding with a plankton substitute on a regular basis, and can, therefore, be recom­mended only to those willing to expend the extra effort.

Following is a list of stony coral genera available to saltwater aquarium hobbyists. I have arranged the genera by families, to facili­tate discussion of their requirements, which tend to remain constant within families.

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            Stony Coral
  • ACROPORIDS AND POCILLOPORIDS
      These large families of stony corals, com­prised of the genera Acropora, Montipora, Anacropora, Astraeopora, Pocillopora, Seriatopora, Stylophora, Palauastrea, and Madracis, include most of the branching stony coral genera available in the saltwater aquarium trade.Although exacting in their requirements, rapid growth rates and the ease with which many species can be propagated from cuttings have resulted in increasing popularity.
  • PORITIDS
      Goniopora, Flowerpot Coral or Sunflower Coral, is considered difficult to maintain. In addition, it is easily damaged by careless han­dling. A related genus, Alveopora, is almost indistinguishable from Goniopora, and is some­times imported.
          Several species in the large genus Porites are imported for the saltwater aquarium these days, with branching types, such as P cylindrica, being popular. They do not appear to be as amenable to saltwater aquarium conditions as some other corals, but this may perhaps be due to collecting and ship­ping damage, rather than to any inherent deli­cacy of the genus.
  • FUNGIIDS
      Heliofungia actiniformis, Plate Coral, needs a soft, sandy substrate and plenty of room. It is not really suitable for the typical reef tank, but is spectacular in a tank designed with its special needs in mind. Fungiids are the only commonly available corals capable of moving from place to place.
          Herpolitha limax, Slipper Coral or Hedge­hog Coral is a close relative of Heliofungia and needs similar conditions. Both these species are shallow water corals. Give them intense light and moderate, not forceful, current.
  • FAVIIDS
      Favia and Favites are both called Closed Brain Coral. Several kinds are imported. Make sure any specimen you select is a complete colony, not just a chunk hacked from a larger specimen. In the latter case, the saltwater aquarium lifes­pan of the specimen may be limited. Other favi­ids that are available to aquarists include Caulastrea, Goniastrea, Platygyra, Leptoria, and Montastrea. 
  • TRACHYPHYLLIIDS
      Trachyphyllia geoffreyi, Open Brain Coral, is a hardy and attractive species that is regularly imported. This coral is a single, large polyp. The skeleton is in the form of an inverted cone, attached at the apex to a hard substrate when young, later breaking off to sit in soft sand or mud as an adult. As a result of this growth form, the coral is easily removed without damage, which may explain why aquar­ium specimens do so well. The related genus Wellsophyllia is also imported for the saltwater aquarium.
  • CARYOPHYLLIIDS
      Several species of Euphyllia are available to aquarists, and all make good additions to the reef tank. All have relatively long tentacles, and must not be placed close to other invertebrates, which they may sting.
      Euphyllia ancora, Anchor Coral, Ham­merhead Coral, Hammer Coral or Ridge Cor­al, has a curved extension at the end of each tentacle, giving the appearance of little ham­mers or anchors. Very desirable as an aquar­ium specimen, this Euphyllia should not be purchased if it hows signs of brown, slimy material forming at the margins of the colony, an indication that the coral has been attacked by a protozoan Helicostoma. Sometimes, a two minute dip in fresh water will eliminate this pest, which apparently gains a foothold when the coral is damaged by rough handling. Euphyllia species seem particularly prone to this problem, although other corals are suscep­tible, as well.
      Euphyllia divisa, Wall Coral, Frogspawn Coral or Vase Coral, gets its colorful common name from the appearance of the tentacles. They sport numerous tubercles and white spots, suggesting a mass of frog's eggs when viewed at a distance. It is another good saltwater aquarium species, although, again, subject to protozoan attack.
      Torch Coral, Euphyllia glabrescens, looks like a cluster of ice cream cones attached at their apices, with long tentacles streaming up from where the ice cream should be, suggesting an Olympic torch. Catalaphyllia jardiniei, Elegance Coral, is closely related to Euphyllia. This is one of the most popular, hardy and spectacular stony corals. Like Trachyphyllia, this species is easy to collect in undamaged condition. In addition, the polyps can withdraw completely into the skeleton, where they are adequately protected during transport, and the coral rarely arrives at its destination in damaged condition. It is some-what restricted in natural distribution, however, and commands a premium price. If I were only going to have one stony coral, though, this would be my choice. Specimens can double in size in six months time.
      Bubble corals are also caryophyllids. Plerogyra sinuosa, Common Bubble Coral, may be pale blue, brownish or greenish in color, as well as creamy white. This is a commonly avail­able and popular species. Beware of damaged specimens, which may not recover in the aquar­ium. Physogyra, which looks very similar, is called Pearl Bubble Coral or Octopus Coral. All caryophylliid corals can be fed small pieces of shrimp or fish, which will enhance their rate of growth. They may also eat unwary gobies or other small fish. Do not, 'however, make the mistake of overfeeding, or feeding too frequently, and bear in mind that giving no food at all does no harm. All these species rely heav­ily upon their zooxanthellae, and can 'survive and grow with this food source alone.
  • DENDROPHYLLIDS
      Turbinaria turbinata, Chalice Coral, Cup Coral or Wineglass Coral, is so named because the skeleton is shaped like a goblet with a fat stem by which the coral is attached to a hard substrate. Thin, brownish tissue covers the entire surface, so it is important to get a speci­men still attached to a small piece of substrate, not one that has merely been snapped off above the point of attachment. The large, flowerlike polyps are borne only on the inside of the "gob­let." Turbinaria is interesting in appearance, and easy to keep. A related, plate-like form is also imported, and is frequently the color of deli mustard. It also makes an attractive addition to the saltwater aquarium. Tubastrea, a non-photosynthetic stony coral imported for its bright orange coloration, is a dendrophyllid.

  • OTHER STONY CORALS
      Many desirable species of stony corals are less frequently encountered than those just dis­cussed. Family Agaricidae (Pavona, Leptoseris. Pachyseris), Family Oculinidae (Galaxea), Family Pectinidae (Pectinia), Family Mussidae(Blastomussa, Cynarina, Scolymia, Symphyllia, Lobophyllia), Family Merulinidae (Hydnophora), are all regularly collected, and one may even find other genera offered from time to time. There are also a few species of coe­lenterates that produce a calcified skeleton that are not true stony corals, even though they are generally regarded as such by people in the saltwater aquarium business. Since they are sometimes imported for the saltwater aquarium, I mention them here. Tubipora is a stoloniferan soft coral, and is discussed below. Heliopora, Blue Coral, is fre­quently seen as a dead skeleton in saltwater aquarium shops, and is collected because of the pretty blue color. The living coral is rather drab, and is sometimes found on live rock. This species is sufficiently different from other coelenterates to warrant placement in its own order, Coenothecalia, within the Octocorallia. ,Heliopora is easily confused with Millepora, Fire Coral, a hydrozoan (Class Hydrozoa) that is found throughout the tropics, and often shows up on live rock specimens. Contact with this coral can cause a painful rash. Despite this drawback, it is a good saltwater aquarium species. Also placed in a separate order (Stylasterina) apart from other hexacorals is the lovely, delicate Stylaster. The colorful, arborescent colonies are found under overhangs. As with other non-photosynthetic coelenterates, Stylaster, though beautiful, is not for beginners.
  • Soft Coral
  • OCTOCORALS
      There are two large groups of "soft corals," which differ from the other coelenterates in that their tentacles are always produced in multi­ples of eight. Thus, the collective term used biologists for this group of coelenterates is "octo­corals." If you feel like counting tentacles, you can verify this fact for yourself. The tentacles of all octocorals also differ in structure from those of the hexacorals, in that they often have tiny branches along them that impart a feathery appearance. Biologists divide this group into two subdivisions, and this is how we will consid­er them here. Classification of the subgroups is based upon the structure of the supporting skeleton, which in one group is stiffened, and in the other, is composed of loose pieces of skeletal material.
  • GORGONIANS
      Soft corals with a stiffened skeleton are gorgonians, commonly called sea fans and sea whips. There are a great many species or these, and they come in a wide variety of colors. Sea whips are branched, like a leafless tree, while in sea fans the branches are fused together, forming a network or mesh, resembling window screening. In all cases, tiny polyps are arrayed along the branches; each gorgonian typically has hundreds of polyps. The color of the polyps, and sometimes the color of the skeleton, is an indication of the light requirements for gorgon­ians. Those in which the polyps are brown or green in color, require light; those with white, red or yellow polyps, generally prefer shaded locations. Specimens in which the skeleton is red, orange or yellow are also generally found in the shade; brownish, blue or purple skeletons typically belong to species that live out in the open sun. Accurately determining the species of a particular gorgonian specimen may be quite difficult, but this should not matter to the aquarist, since all gorgonians have the same basic requirements. Provide light for those that require it. In all cases, provide good water movement. Strong currents seem to be very important for these organisms. With few excep­tions, gorgonians collected for the saltwater aquarium originate in Florida, the Caribbean, or the Gulf of Mexico. The non-photosynthetic species range from easy to impossible, in terms of their saltwater aquarium care, and should only be attempted by aquarists who have gained some experience.
  • ALCYONARIANS
      Soft corals with "flabby" skeletons com­posed of loose aggregations of spicules are clas­sified as "alcyonarians." In Florida and the Caribbean gorgonians dominate shallow water hard bottom habitats; in the Indo-Pacific this ecological niche is occupied by the alcyonarians. Several of these are commonly imported for the saltwater aquarium.
  • LEATHER CORALS
      Sarcophyton, usually called' Leather Mushroom Soft Coral, is among the most suit­able of alcyonarians for a reef tank. In all of the leather corals, the body mass is brownish or yel­lowish in color, and the polyps are embedded in this skeletal mass. Leather corals prefer high light intensities. They often grow almost up to the surface of the ocean in nature. It is not unusual for the polyps of leather corals to remain contracted for several days after a change in water conditions, such as moving them from one saltwater aquarium to the other.
      Colt Soft Coral (Cladiella) is branched like a tree. It is brown in color, with the "trunk" usually much lighter than the branches. This lovely species has requirements similar to Sarcophyton. Other genera of leather corals regularly seen in saltwater aquarium shops include Litophyton, Lobophytum, and Sinularia. All of the photosynthetic alcyonarians can be propa­gated from cuttings taken by various methods, and are beginning to form the foundation for small marine aquaculture operations all over the United States.
  • STOLONIFERANS
      Starburst Soft Coral, also called Green Star Polyps, is frequently misidentified as Organ Pipe Coral (Tubipora musica), a closely related species with a calcified skeleton. In Starburst Soft Coral, the skeleton is a rubbery flattened sheet that encrusts a solid substrate Each polyp resides in a short tube that projects upward from the basal sheet . The polyps them. selves are generally pale green with bright green centers, or are an overall lime green color When expanded, this is a beautiful species. It also, happily, one of the hardiest and moss durable of reef tank organisms, and can be highly recommended even to the beginner. Its only light, under which it will grow and spread. It will even tolerate dim light, but in this situation it never looks at its best.
  • XENIIDS
      Several species of soft corals hold special interest for reef enthusiasts because they exhib­it continuous, pulsing movements. These are col­lectively referred to as "pulse polyps," Family Xeniidae, and may be species of either Xenia or Anthelia. In the former, the columns of the indi­vidual polyps are long and thin, and the, crown of tentacles reminds one of a daisy. They are brownish in color. In Anthelia, the feathery, white polyps are attached at the base to form a cluster anchored to a rock or other hard surface. Both Xenia and Anthelia are hardy and repro­duce themselves readily in the saltwater aquarium. Other alcyonarians are imported for the saltwater aquarium. Most will require the same lighting and water conditions that leather corals need. One exception is Dendronephyta, which prefers shade, and does not contain zooxanthellae. It is very challenging to keep and can only be recommended to advanced aquarists capable of providing for its rigorous requirements, includ­ing live phytoplankton.
  • AI PTASIA
      One coelenterate should be discussed here because it is frequently a pest. Many authors recommend eliminating the anemone Aiptasia whenever possible, citing its ability to "nettle'. other coelenterates. I have heard a number of recommendations for eliminating Aiptasia, None, alas, are really practical. For example, you can inject each individual anemone with potassium, sodium, or calcium hydroxide. This will certainly do them in, but is an extremely tedious undertaking. Pulling or scraping their off the glass and rocks does no good, since ever a tiny piece of the anemone's basal disk wil regenerate an entire new individual in a few weeks time. Other suggestions include intro ducing a Copperband or Raccoon butterflyfish One waits until all the Aiptasia have been eaten, and then attempts to devise a way to catch the fish agail before it starts feeding on coral polyps. I am told tha German aquarists at one time actually set uptanks and deliberately cultivated abundant Aiptasia for use as "training camps" for imma­ture Copperband Butterflies destined for duty as Aiptasia removers in the reef tank. The the­ory behind this is that the butterflyfish, having learned to feed only on Aiptasia, will continue this practice once moved from the "training camp" to the front lines. This seems to me like a lot of trouble for an approach that -may not always work. Further, if it does not work there is the problem of extracting the agile butterfly-fish from the carefully arranged reef tank, with all those crannies and caves. Enthusiasm dims at the prospect of trying to catch any fish, much less a butterflyfish, under such circumstances. The common, cheap little Dancing, Shrimp, Rhyncocinetes uritai, will do the job, although you have to worry about them starting in on your Elegance Coral after the Aiptasia are all gone. Aiptasia are usually introduced with live rock, and seem to flourish best in tanks that are too warm, and which have a heavy nutrient load. The best approach to eliminating them is to create conditions more favorable to desirable Species such as stony corals, i.e., cool, nutrient-poor water.

           One final note about corals. Handle them carefully. They are easy to damage, and damage can lead to fatal bacterial infections. Make sure you understand the requirements of stony corals before you attempt to keep them in the saltwater saltwater aquarium. Learn reef keeping techniques with the easiest species first. Stony corals are among the most beautiful and fragile of the reef's trea­sures. If you decide to keep them in the aquari­um, treat them with the care and respect they deserve. Fortunately, in recent years there have been great strides in captive propagation of many stony coral species, and the future looks bright for continued enjoyment of them in hob­byist saltwater aquarium.

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